from The American Heritage® Dictionary of the English Language, 4th Edition

  • adj. Walking with the long axis of the body parallel to the ground. Used of quadrupeds.

from The Century Dictionary and Cyclopedia

  • Carrying the body in a horizontal position, as do the majority of quadrupeds: opposed to orthograde.


Latin prōnus, leaning forward; see prone + -gradus, walking (from gradī, to move; see retrograde).
(American Heritage® Dictionary of the English Language, Fourth Edition)


  • Without interfering with the ape, the researchers used a forceplate to record the forces exerted by the gibbon's feet as it leapt, and two high speed cameras to record details of its body position, and the angles of its legs, body and arms. his revealed that the gibbon uses four distinct modes of leaping, categorised as the orthograde single footed take-off, the orthograde two footed take-off, the orthograde squat jump and the pronograde single footed take off.

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