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The ecoclimate of this ecoregion ranges from high and mid-boreal and perhumid mid-boreal to Oceanic, Atlantic, and maritime mid-boreal.
Climatically, the ecoregion is characterized by a perhumid sub-alpine bioclimate (over 1,800 millimeters (mm) of annual rainfall; average annual temperature of about 3-5ºC).
The current climate of these mountains is much wetter than the surrounding lands, with perhumid (rain every month) conditions and rainfall up to 3,000 millimeters (mm) per year recorded in the eastern Uluguru Mountains.
Trees in arid environments are innately well adapted to cope with wide swings in temperature versus their perhumid counterparts, hence, the prevalence of evergreen as opposed to deciduous types.
Humid and perhumid continental areas a couple of hundred to a hundred miles south of the the northern limits of tree growth may also be candidates, but I’d be concerned about soil freeze thaw variation causing distortion.