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While it can't be entirely ruled out that large fishes (remember that there were some very big fishes in the Jurassic seas of Europe) made some of traces, ichthyosaurs and plesiosaurs, and perhaps thalattosuchian crocodilians, are the most likely candidates.
Marine reptiles are what make the Oxford Clay really interesting, and they’re represented by three groups: thalattosuchian crocodilians, plesiosaurs and ichthyosaurs.